Background: T-cells can be genetically modified to express artificial Chimeric Antigen Receptors (CARs) enabling them to target and lyse cancer cells. CAR T-cells targeting the B-cell antigen, CD19, have induced complete remissions in >90% of patients with relapsed/refractory Acute Lymphoblastic Leukemia and >50% in relapsed/refractory Diffuse Large B-cell Lymphoma. Currently, CAR T-cells are produced with random gene integrating techniques to introduce the CAR transgene. Additionally, they are manufactured individually for each patient from autologous T-cells which is time-consuming and expensive. To mitigate these problems, off-the-shelf CAR T-cells can be produced from a healthy donor. However, their efficacy may be limited by recipient T-cell mediated rejection of mismatched human leukocyte antigens (HLA), while they may also cause toxicity in the form of graft-versus-host disease (GVHD) via their native T-cell receptor (TCR).
Aims: To prevent GVHD, we generated off-the-shelf CAR T-cells by CRISPR/Cas9 knockout of the TCR complex. Additionally, we investigated knockout of various HLA Class I and II-associated gene targets, to prevent CAR T-cell rejection.
Methods: T-cells from healthy donors were transfected by electroporation with Cas9 enzyme, single guide RNA and CAR19 transgene, expanded for 3 weeks, and stimulated with interleukin-15 and irradiated autologous peripheral blood mononuclear cells.
Results: CRISPR-edited CAR T-cells exhibited comparable expansion, memory phenotypes and expression of immunoinhibitory molecules to our Phase I clinical trial CAR T-cells produced with the non-specific Super PiggyBac transposon system. In screening potential HLA-associated targets for HLA downregulation, we discovered a single novel target gene that efficiently downregulates HLA Class I and II expression on CAR T-cells.
Translational significance: Our data demonstrates that CRISPR/Cas9 directed against novel targets can be used to efficiently generate CAR T-cells lacking TCR and HLA. This developmental research will lay the crucial groundwork for the development of future off-the-shelf T-cell therapy products to target various malignancies and infections.